Anther consists of microsporangia or pollen sacs. The archesporium gives rise to parietal cells and primary sporogenous tissue. Sporogenous cells divide to form pollen grain or microspore mother cells. They are diploid and connected by plasmodesmata. The microspore, mother cells consists of a callose wall inner to the cell wall. The mother cell then undergoes meiosis and forms tetrads of microspores. Finally the wall of the mother cell degenerates and pollen grains are separated.